Rome attacked the thin Carthaginian centre, which gave way on Hamilcar's orders. The first and second Roman squadrons pursued, opening a gap because the third was towing the horse transports and the fourth had to remain close to protect them. Once the separation was sufficient, a signal from Hamilcar brought his ships around together to fight their pursuers. Carthaginian speed allowed repeated attacks and rapid disengagement. Roman boarding aggression and the corvus offered a different advantage whenever an enemy came close, and both consuls took part in hand-to-hand combat. The separation also gave Hanno room to begin his attack on the rear guard.
Notable Point · 1:01:34-1:02:52
How the Carthaginian retreat divided Rome's fleet
Towing the horse transports kept Rome's rear squadrons from following the pursuit, allowing Carthage to divide the battle.